Journal:

2018. 26 (1)

Publicatione: 
REVIEW OF MITES (ACARI) ASSOCIATED WITH THE EUROPEAN SPRUCE BARK BEETLE, IPS TYPOGRAPHUS (COLEOPTERA: CURCULIONIDAE: SCOLYTINAE) IN ASIAN RUSSIA



About authors:

Khaustov A. A., Tyumen State University, Tyumen, Russia
Klimov P. B., University of Michigan, Museum of Zoology, 1109 Geddes Ave., Ann Arbor, Michigan 48109-1079 USA
Trach V.A., I. I. Mechnikov Odessa National University, Shampanskij al. 2, Odessa, 65058 Ukraine
Bobylev A.N., Tyumen State University, Tyumen, Russia
Salavatulin V.M., Tyumen State University, Tyumen, Russia; State Agrarian University of the Northern Trans-Urals, Tyumen, Russia
Khaustov V. A., Tyumen State University, Tyumen, Russia
Tolstikov A.V., Tyumen State University, Department of Zoology and Evolutionary Ecology of Animals, 10 Semakova str., Tyumen, 625003 Russia

Annotation:

A total of 68 mite species were collected on Ips typographus and in their galleries in Siberia and in the Far East of Russia: Cercoleipus kuznetsovi (Cercomegistidae), Pleuronectocelaeno barbara, P. japonica (Celaenopsidae), Uroobovella ipidis (Urodinychidae), Trichouropoda polytricha, T. karawaiewi, T. structura (Trematuridae), Schizosthetus sp. (Parasitidae), Dendrolaelaps (Ipidodendrolaelaps) quadrisetus, Dendrolaelaps (Insectolaelaps) pini, Dendrolaelaps (Apophyseodendrolaelaps) disetosimilis (Digamasellidae), Macrocheles (s. str.) ivanovi (Macrochelidae), Gamasellodes tatricus (Ascidae), Proctolaelaps fiseri, P. hystricoides, P. juradeus (Melicharidae), Ereynetes (Huntereynetes) sp. (Ereynetidae), Mexecheles virginiensis (Cheyletidae), Dolichocybe sibiriensis (Dolichocybidae), Pyemotes sp. (Pyemotidae), Aethiophenax ipidarius (Acarophenacidae), Microdispodides moseri, Elattoma crossi (Pygmephoridae), Coleopygmephorus bogenschutzi, C. hylophilus, C. typographus (Neopygmephoridae), Scutacarus pseudoplurisetus, S. scolyti, S. muscocicola, S. palustris (Scutacaridae), Iponemus gaebleri gaebleri, Tarsonemus striatus, T. (Schaarschmidtia) bistridentati, T. typographi, Heterotarsonemus lindquisti (Tarsonemidae), Siculobata (Paraleius) leontonycha (Scheloribatidae), Acarus sp., Boletoglyphus boletophagi, Forcellinia wasmanni, Ipsoglyphus bochkovi, Kuzinia laevis, Naiadacarus sp., Schwiebea sp. eurynympha-group [punctate dorsum], Sch. sp. eurynympha-group [small, unsclerotized, 1a, 3a filiform], Sch. nesbitti, Sch. nova, Sch. sp.1 nova-group [two tibial setae, regular attachment organ], Sch. sp. nova-group [two tibial setae, wide attachment organ], Sch. sp. nova-group [granulate dorsum, short legs III–IV], Sch. scheucherae, Thyreophagus corticalis, Th. entomophagus, Th. aff. odyneri, Tyrophagus putrescentiae (Acaridae), Divilia aff. occidentalis, D. sp., Hemisarcoptes sp., Nanacarus sp.1, N. sp.2, Superioropus huronmontanus (Hemisarcoptidae), Parawinterschmidtia kneissli, Saproglyphus aff. hagensis, S. waldorfae (Winterschmidtiidae), Histiostoma ovalis, H. piceae, H. pini, Probonomoia pini, P. sibirica (Histiostomatidae). The following taxa are recorded for the first time: for the Palaearctic—Superioropus huronmontanus (genus and species); for Russia—Pleuronectocelaeno japonica, Trichouropoda karawaiewi, Dendrolaelaps pini, Gamasellodes tatricus, Scutacarus palustris, Tarsonemus typographi, Heterotarsonemus lindquisti, Hemisarcoptes, Saproglyphus waldorfae, Naiadacarus, Schwiebea nesbitti, Histiostoma pini, and H. ovalis. Of them, 60 species were collected by catching beetles with pheromone traps and examining them for phoretic mites.
We provide information on the current distribution, host association, and biology of these mites. In this article, we also discuss the following information, which we have documented: our biological observations of mites in the beetles’ galleries, abundance of phoretic mites on hosts, as well as specific sites where they occur. To facilitate morphological diagnostics, common mite species were illustrated using a Scanning Electron Microscope (SEM), as well as light microscopy.

DOI: 10.21684/0132-8077-2018-26-1-3-79

Bibliography:

Ahadiyat, A. and Akrami, M.A. 2015. Oribatid mites (Acari: Oribatida) associated with bark beetles (Coleoptera: Curculionidae: Scolytinae) in Iran, with a review on Paraleius leontonychus (Berlese) and a list of bark beetles in association with this species. Persian Journal of Acarology, 4: 355–371.
Ahadiyat, A., Khaustov, A.A., Seeman, O.D. and Valizadeh, A. 2011. First record of mites of the genus Cercoleipus Kinn, 1970 (Mesostigmata: Trigynaspida: Cercomegistidae) from Iran. In: S. Kazemi and A. Saboori (Eds.). Abstract and Proceeding Book of the First Persian Congress of Acarology, Kerman, Iran, p. 2.
Ahadiyat, A., Ostovan, H., Gwiazdowicz, D. and Valizadeh, A. 2012. Celaenopsid mites (Mesostigmata) associated with the Mediterranean pine engraver beetle, Orthotomicus erosus (Wollaston) (Coleoptera: Curculionidae: Scolytinae), in Iran. In: A. Sarafrazi, M. R. Asef, M. Mozhdehi, M. Mozh-
dehi, S. Solhjouy Fard and T. Abdollahi (Eds.). Proceedings of the 20th Iranian Plant Protection Congress. 26–29 August 2012. Shiraz University, Iran, p. 475.
Al-Atawi, F., Klompen, H. and Moser, J.C. 2002. Redescription of Schizosthetus lyriformis (McGraw and Farrier, 1969) (Parasitiformes: Parasitidae), with revision of the genus. International Journal of Acarology, 28: 341–360.
Alekseev, G.A., Marchenko, I.I., Savvinov, G.N. and Boeskorov, V.S. 2011. Soil gamasid mites (Acari: Mesostigmata) of Central Yakutia. In: V.E. Kipyatkov and D.L. Musolin (Eds.). Materials of the International Scientific Conference “Fundamental Problems of Entomology in the 21st Century” (16–20 May 2011, Saint Petersburg, Russia). St. Petersburg University, Saint Petersburg, p. 7.
André, M. 1942. Sur lʼHemisarcoptes malus Shimer (=coccisugus Lignières). Bulletin du Muséum national dʼhistoire naturelle. 2ème Série, 14: 173–180.
Arlian, L.G., Geis, D.P., Vyszenski-Moher, D.L., Bernstein, I.L. and Gallagher, J.S. 1984. Antigenic and allergenic properties of the storage mite Tyrophagus putrescentiae. Journal of Allergy and Clinical Immunology, 74: 166–171.
Athias-Henriot, C. 1959. Contribution to the genera Podocinum Berlese and Pleuronectocelaeno Vitzthum in Algeria and notes on the genus Aceosejus Sellnick and Seiodes Berlese (Mesostigmata). Acarologia, 1: 24–38.
Athias-Henriot, C. 1982. Schizosthetus n.g. (type Eugamasus lyriformis McGr. & Farr., 1969) avec deux espèces nouvelles (Parasitiformes, Parasitidae). Acarologia, 23: 207–214.
Baker, E.W. 1949. A review of the mites of the family Cheyletidae in the United States National Museum. Proceedings of the United States National Museum, 99: 267–320.
Balazy, S. and Kiełczewski, B. 1965. Tarsonemoides gaebleri Schaarschmidt (Acar. Tarsonemidae) jajozerny roztocz w zerowiskach kornika drukarza Ips typographus (L.). Polskie Pismo Entomologiczne B, 37/38: 7–18.
Bayoumi, B.M. and Al-Khalifa, M.S. 1983. Soil mites and Collembola from Al-Quasim Province, Saudi Arabia. Arab Gulf Journal of Scientific Research, 1: 179–186.
Berlese, A. 1920 (1921). Centuria quinta di Acari nuovi. Redia, 14: 143–195.
Berlese, A. 1903 (1904). Acari nuovi. Manip. II. Redia, 1: 258–280.
Bochkov, A.V. and Abramov, V.V. 2016. To fauna of the free-living Cheyletidae (Acariformes: Cheyletoidea) of the European part of Russia. Systema-
tic and Applied Acarology, 21: 335–346.
Brasier, C.M. 1978. Mites and reproduction in Ceratocystis ulmi and other fungi. Transactions of the British Mycological Society, 70: 81–89.
Bregetova, N.G. 1977a. [Family Aceosejidae]. In: M.S. Gi-
larov and N.G. Bregetova (Eds.). Opredelitel’ obytayshchikh v pochve kleshchey Mesostigmata [Identification key to soil-inhabiting mites Mesostigmata]. Nauka, Leningrad, pp. 169–226.
Bregetova, N.G. 1977b. [Family Macrochelidae]. In: M.S. Gilarov and N.G. Bregetova (Eds.). Opredelitel’ obytayshchikh v pochve kleshchey Meso-
stigmata [Identification key to soil-inhabiting mites Mesostigmata]. Nauka, Leningrad, pp. 346–411.
Bregetova, N.G. and Koroleva, E.V. 1960. The macrochelid mites (Gamasoidea, Macrochelidae) of the USSR. Parazitologicheskiy Sbornik, 19: 32–154.
Bugrov, S.A. 1995. [New species of the genus Schwiebea (Astigmata, Acaridae) from Russia and adjacent countries]. Zoologicheskii Zhurnal, 74: 61–75.
Bugrov, S.A. 1997. [Free-living Astigmata (Acariformes) of the Moscow District fauna]. Zoolo-
gicheskii Zhurnal, 76: 147–156.
Burjanadze, M., Moser, J.C., Zimmermann, G. and Kleespies, R.G. 2008. Antagonists of the spruce bark beetle Ips typographus L. (Coleoptera: Scolytidae) of German and Georgian populations. IOBC-WPRS Bulletin, 31: 245–250.
Castagnoli, M., Liguori, M. and Nannelli, R. 1994. Mite communities associated with chestnut blight. Atti del Congresso Nazionale Italiano di Entomologia, 17: 465–468.
Čejka, M. and Holuša, J. 2014. Phoretic mites in uni- and bivoltine populations of Ips typographus: a 1-year case study. Turkish Journal of Zoology, 38: 569–574.
Černý, V. and Samšiňák, K. 1971. Nadkohorta Acaridiae. In: M. Daniel and V. Černý (Eds.). Klíč zvířeny ČSR. Želvušky, jazyčnatky, klepítkatci, sekáči, pavouci, štírci, roztoči. Československá Akademie Věd, Praha, pp. 496–529.
Chaires-Grijalva, M.P., Estrada-Venegas, E.G., Equihua-Martinez, A., Moser, J.C., Sanchez-Martinez, G., Vazquez-Rojas, I.M. et al. 2013. Mesostigmados (Acari: Mesostigmata) asociados con Dendroctonus rhizophagus de Chihuahua, Mexico. Revista Mexicana de Biodiversidad, 84: 1235–1242.
Chant, D.A. 1963. The subfamily Blattisocinae Garman (=Aceosejinae Evans) (Acarina: Blattisocidae Garman) (=Aceosejidae Baker & Wharton) in North America, with descriptions of new species. Canadian Journal of Zoology, 41: 243–305.
Chmielewski, W. 1969. Obserwacje nad biologią nowego dla akarofauny polskiej gatunku Kuzinia laevis (Dujardin, 1849) (Acarina, Acaridae) [=Observations on the biology of the species Kuzinia laevis (Dujardin, 1849) (Acarina, Acaridae), new in Polish acarofauna]. Polskie Pismo Entomologiczne. Bulletin Entomologique de Pologne, 39: 603–617.
Chmielewski, W. 1975. Z badań nad Thyreophagus entomophagus (Laboulbène). Zeszyty Problemowe Postępów Nauk Rolniczych, 171: 253–259.
Chmielewski, W. 1977. Wyniki obserwacji powiazan roztoczy z owadami (Acari-Insecta). Polskie Pismo Entomologiczne, 47: 59–78.
Chmielewski, W. 1991. Stored products mites (Acaroidea) in Polish bee hives. In: Modern acarology. Volume I: proceedings of the 8 International Congress of Acarology, held in Ceske Budejovice, Czechoslovakia, 6–11 August 1990. SPB Academic Publishing, The Hague, The Netherlands, pp. 615–619.
Cooreman, J. 1963. Notes et observations sur quelques Acariens inféodés aux Coléoptères Scolytides de la faune belge. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique—Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, 39: 1–48.
Cotter, M., Siebers, R., Pike, A., Fitzharris, P. and Crane, J. 2011. Storage mites in flour samples in Wellington, New Zealand. Journal of Investigational Allergology and Clinical Immunology, 21: 410–411.
Cross, E.A. and Moser, J.C. 1971. Taxonomy and biology of some Pyemotidae (Acarina: Tarsonemoidea) inhabiting bark beetle galleries in North American conifers. Acarologia, 13: 47–64.
Cross, E.A., Moser, J.C. and Rack, G. 1981. Some new forms of Pyemotes (Acarina: Pyemotidae) from forest insects, with remarks on polymorphism. International Journal of Acarology, 7: 179–196.
Davydova, M.S. and Nikolskii, V.V. 1986. Gamazovye kleshchi Zapadnoi Sibiri. [Gamasid mites of Western Siberia.]. Novosibirsk, Nauka. 1–123 pp.
De Camargo Barbosa, M.F., OConnor, B.M. and De Moraes, G.J. 2016. A new species of Thyreophagus (Acari: Acaridae) from Brazil, with notes on species associated with stored food and human habitats and a key to species of this genus. Zootaxa, 4088: 279–291.
de la Torre Morin, F., de la Coba, R.A., Gallego, M., Iraola, V. and Fernandez-Caldas, E. 2006. Sensitization to Thyreophagus entomophagus and other mite species in Cuban patients with allergic asthma and/or rhinoconjunctivitis. Journal of Allergy and Clinical Immunology, 117: S119–S119.
De Moraes, G.J., Britto, E.P.J., Mineiro, J.L.D.C. and Halliday, B. 2016. Catalogue of the mite families Ascidae Voigts & Oudemans, Blattisociidae Garman and Melicharidae Hirschmann (Acari: Mesostigmata). Zootaxa, 4112: 1–276.
Dobrev, D. and Mahunka, S. 1991. Data to the scutacarid fauna of the Bátorliget nature conservation areas (Acari: Heterostigmata). In: The Bátorliget Nature Reserves—after forty years. Hungarian Natural History Museum, Budapest, pp. 719–726.
Domrow, R. 1992. Acari Astigmata (excluding feather mites) parasitic on Australian vertebrates: an annotated checklist, keys and bibliography. Invertebrate Taxonomy, 6: 1459–1606.
Dzibladze, K. N. 1969. A new species of predatory mite (Hemisarcoptidae) attacking the orange scale, Cornuaspis beckii Newman (Homoptera, Coccoidea), in Western Georgia. Entomologicheskoe Obozrenie, 48: 435–436.
Elkawas, H. 2011. Acarines As Biological Control Agents: An Overview of Bio-relationships between Mites and Insects in Egypt LAP LAMBERT Academic Publishing. 1-128 pp.
Erban, T., Rybanska, D., Harant, K., Hortova, B. and Hubert, J. 2016. Feces derived allergens of Tyrophagus putrescentiae reared on dried dog food and evidence of the strong nutritional interaction between the mite and Bacillus cereus producing protease bacillolysins and exo-chitinases. Frontiers in Physiology, 7.
Ermilov, S.G. and Mokrousov, M.V. 2008. Acariform mites (Acariformes) phoresing on capricorn beetles (Coleoptera, Cerambycidae). Povolzhskii Ekologicheskii Zhurnal, 3: 222–225.
Ermilov, S.G. and Mokrousov, M.V. 2010. Mites phoretic on wasps (Hymenoptera: Ampulicidae, Sphecidae, Crabronidae) in the middle Volga region. Entomoligicheskie i parazitplogicheskie issledovaniya v Povolzh’e, 8: 85–95.
Ermilov, S.G., Mokrousov, M.V. and Dmitrieva, I.N. 2008. Snout beetle acarofauna (Coleoptera, Curculionoidea). Povolzhskii Ekologicheskii Zhurnal, 1: 65–68.
Ermilov, S.G., Mokrousov, M.V. and Mukhanov, A.V. 2006. Phoretic associations between acariform mites (Acariformes) and click beetles (Coleoptera, Elateridae). Povolzhskii Ekologicheskii Zhurnal, 2/3: 176–179.
Ermilov, S.G., Mokrousov, M.V. and Mukhanov, A.V. 2007. Acarofauna of bark beetles (Coleoptera, Scolytidae) in the Nizhniy Novgorod region. Povolzhskiy Journal of Ecology, 1: 67–70.
Evans, G.O. 1958. A revision of the British Aceosejinae (Acarina: Mesostigmata). Proceedings of the Zoological Society of London, 131: 177–229.
Fain, A. 1972. Notes sur les hypopes des Saproglyphidae (Acarina: Sarcoptiformes) 2. Redefinition des genres. Acarologia, 14: 225–249.
Fain, A. 1974. Observations sur la faune acarologique du sol dans une savane de Côte dʼIvoire (Acaridae: Sarcoptiformes). Bulletin de lʼInstitut Fondamental de lʼAfrique Noire Série A, 36: 69–91.
Fain, A. 1977. Notes sur le genre Schwiebea Oudemans, 1916 (Acarina, Astigmata, Acaridae). Bulletin & Annales de la Societe Royale Belge dʼEntomologie, 113: 251–276.
Fain, A. 1982. Revision des genres Thyreophagus Rondani, 1874 et Michaelopus Fain & Johnston, 1974 (Acari, Acaridae) avec description de neuf especes nouvelles. Bulletin de lʼInstitut Royal des Sciences Naturelles de Belgique Entomologie, 54: 1–47.
Fain, A. 1987. Observations on the hypopi of the genus Forcellinia Oudemans, 1924 (Acari, Acaridae). Bulletin de lʼInstitut Royal des Sciences Naturelles de Belgique Entomologie, 57: 111–120.
Fain, A. 1988. Observations on Congovidia Fain & Elsen, 1971 and allied genera (Acari, Hemisarcoptidae). Bulletin & Annales de la Societe Royale Belge dʼEntomologie, 124: 125–130.
Fain, A. and Baugnee, J.Y. 1996. Acariens phorétiques ou parasites récoltés sur des insectes du sud de la Belgique. Deuxième note. Bulletin et Annales de la Societe Royale Belge dʼEntomologie, 132: 19–33.
Fain, A. and Beaucournu, J.C. 1972. Notes sur les hypopes vivant en association phorétique sur les puces en France (Acarina: Sarcoptiformes). Acarologia, 13: 522–531.
Fain, A. and Beaucournu, J.C. 1993. Les hypopes des astigmates (Acari) phoretiques sur les puces (Siphonaptera) de mammiferes et dʼoiseaux. Bulletin de lʼInstitut Royal des Sciences Naturelles de Belgique Entomologie, 63: 77–93.
Fain, A. and Greenwood, M.T. 1991. Notes on a small collection of mites (Acari), phoretic on Diptera, mainly Phoridae, from the British Isles. Bulletin de lʼInstitut Royal des Sciences Naturelles de Belgique Entomologie, 61: 193–197.
Fain, A., Hurst, G.D.D., Fassotte, C., Webberley, K.M., Sloggett, J.J. and Majerus, M.E.N. 1997. New observations on the mites of the family Hemisarcoptidae (Acari: Astigmata) phoretic on Coccinellidae (Coleoptera). Bulletin de lʼInstitut Royal des Sciences Naturelles de Belgique Entomologie, 67: 89–94.
Fain, A., Hurst, G.D.D., Tweddle, J.C., Lachlan, R.F., Majerus, M.E.N. and Britt, D.P. 1995. Description and observations of two new species of Hemisarcoptidae from deutonymphs phoretic on Coccinellidae (Coleoptera) in Britain. International Journal of Acarology, 21: 99–106.
Fain, A. and Johnston, D. 1974. Three new species of hypopi phoretic on springtails (Collembola) in England (Acari: Acarididae). Journal of Natural History, 8: 411–420.
Fain, A., Knuelle, W. and Wurst, E. 2000. First description of the hypopial stage of Thyreophagus entomophagus (Laboulbène, 1852) (Acari Acaridae). Bulletin de la Societe Royale Belge dʼEntomologie, 136: 153–156.
Fain, A. and Rack, G. 1987. Allocalvolia habrocytus gen.n., sp.n. (Acari, Winterschmidtiidae) a new hypopus phoretic on Habrocytus elevatus (Walker 1834) (Hymenoptera, Pteromalidae) in Austria. Entomologische Mitteilungen aus dem Zoologischen Staatsinstitut und Zoologischen Museum Hamburg, 8: 309–341.
Fain, A. and Ripka, G. 1998. A new species of Hemisarcoptes Lignières, 1893 (Acari: Hemisarcoptidae) from ornamental trees in Hungary. International Journal of Acarology, 24: 33–39.
Fain, A. and Schwan, T.G. 1984. Three new hypopial nymphs (Acari: Acaridae) phoretic on fleas parasitic on rodents in California, USA. Bulletin & Annales de la Societe Royale Belge dʼEntomologie, 120: 91–97.
Fashing, N.J. 1975. Life history and general biology of Naiadacarus arboricola Fashing, a mite inhabiting water-filled treeholes (Acarina: Acaridae). Journal of Natural History, 9: 413–424.
Fashing, N.J. and Chua, T.H. 2002. Systematics and ecology of Naiadacarus nepenthicola, a new species of Acaridae (Acari: Astigmata) inhabiting the pitchers of Nepenthes bicalcarata Hook. f. in Brunei Darussalam. International Journal of Acarology, 28: 157–167.
Feketeová, Z. 2011. Význam uropodných roztočov (Acari: Uropodina) v ekológii lykožrúta smrekového (Ips typographus). Entomofauna carpathica, 23: 11–19.
Fernández, M., Julio, D. and Moraza, M.L. 2013. Acarofauna associated with Ips sexdentatus in northwest Spain. Scandinavian Journal of Forest Research, 28: 358–362.
Fumouze, A. and Robin, C. 1867. Mémoire anatomique et zoologique sur les Acariens des genres Cheyletus, Glyciphagus et Tyroglyphus. Journal de lʼanatomie et de la physiologie normales et pathologiques de lʼhomme et des animaux, 4: 561–601+Pl. XXII–XXV.
Gerson, U. and Schneider, R. 1981. Laboratory and field studies on the mite Hemisarcoptes coccophagus Meyer (Astigmata: Hemisarcoptidae), a natural enemy of armored scale insects. Acarologia, 22: 199–208.
Goracci, E., Lazzeri, S., Zuccherelli, D., Rossetti, M. and Poggianti, A.M. 1985. [Acari of food storage facilities. An ecologic and immuno-allergic study]. Quaderni Sclavo di diagnostica clinica e di laboratorio, 21: 436–446.
Gorton, C. and Webber, J.F. 2000. Reevaluation of the status of the bluestain fungus and bark beetle associate Ophiostoma minus. Mycologia, 92: 1071–1079.
Griffiths, D.A. 1964. A revision of the genus Acarus L. 1758 (Acaridae, Acarina). Bulletin of the British Museum (Natural History) Zoology, 11: 413–464.
Griffiths, D.A. 1970. A further systematic study of the genus Acarus L., 1758 (Acaridae, Acarina), with a key to species. Bulletin of the British Museum (Natural History) Zoology, 19: 85–118.
Griffiths, D.A., Hodson, A.C. and Christensen, C.M. 1959. Grain storage fungi associated with mites. Journal of Economic Entomology, 52: 514–518.
Grobov, O.F. 1978. Kleshchi medonosnoy pchely (Apis mellifera L.): ikh znachenie i osnovnue printsypy bor’by s kleschchevymi porazheniyami [=Mites of the honeybee (Apis mellifera L.): their significance and main principles of control of diseases caused by mites]. Doctor of Sciences (Habilitation) Thesis. In: All-Union Institute of Experimental Veterinary, All-Union Academy of Agricultural Sciences, Moscow, p. 536.
Gwiazdowicz, D.J. 2007. Ascid Mites (Acari, Mesostigmata) from Selected Forest Ecosystems and Microhabitats in Poland. Poznan, Wydawnictwo Akademii Rolniczej im. Augusta Cieszkowskiego, 2007. 247 pp.
Gwiazdowicz, D.J. 2008. Mesostigmatid mites (Acari) associated with Scolytidae in Poland. In: D.J. Gwiazdowicz (Ed.). Selected problems of acarological research in forests. Wydawnictwo Uniwersytetu Przyrodniczego, Poznan, pp. 59–95.
Gwiazdowicz, D.J., Bloszyk, J. and Gdula, A.K. 2015. Alpha diversity of mesostigmatid mites associated with the bark beetle Ips typographus (L.) in Poland. Insect Conservation and Diversity, 8: 448–455.
Gwiazdowicz, D.J. and Gutowski, J.M. 2012. Records of phoretic mesostigmatid mites (Acari: Mesostigmata) on beetles (Coleoptera: Carabidae, Cerambycidae, Elateridae, Erotylidae, Scolytinae) in the Białowieża Primeval Forest. Polskie pismo entomologiczne, 81: 305–310.
Gwiazdowicz, D.J., Kamczyc, J. and Bloszyk, J. 2011. The diversity of phoretic Mesostigmata on Ips typographus (Coleoptera: Scolytinae) caught in the Karkonosze forest. European Journal of Entomology, 108: 489–491.
Gwiazdowicz, D.J., Kamczyc, J., Teodorowicz, E. and Bloszyk, J. 2012. Mite communities (Acari, Mesostigmata) associated with Ips typographus (Coleoptera, Scolytidae) in managed and natural Norway spruce stands in Central Europe. Central European Journal of Biology, 7: 910–916.
Gwiazdowicz, D.J. and Walter, D.E. 2005. Gamasellodes tatricus sp. nov. (Acari: Ascidae) from Poland. Systematic and Applied Acarology, 10: 61–66.
Haddad Irani Nejad, K., Kamali, K. and Maleki Milani, H. 2001. Mites of Digamasellidae, Rhodacaridae, Phytoseiidae and Ascidae of the cooton fields in Moghan Plain. Agricultural Sciences and Technology, 15 23–33.
Haitlinger, R. 1988. Mites (Acari) occurring on beetles of the genus Carabus Linnaeus, 1758 (Insecta, Coleoptera, Carabidae). Wiadomości Parazytologiczne, 34: 329–346.
Haitlinger, R. 2004. Mites (Acari) occurring on some Coleoptera (Insecta) in Poland. Polskie Pismo Entomologiczne, 73: 3–24.
Haitlinger, R. and Lupicki, D. 2012. Mites (Acari) occurring on some Coleoptera and Diptera in Poland. Zeszyty Naukowe Uniwersytetu Przyrodniczego we Wroclawiu, 587: 17–48.
Halliday, R.B. 2000. Additions and corrections to Mites of Australia: A checklist and bibliography. Australian Journal of Entomology, 39: 233–235.
Halliday, R.B. 2002. Two genera of mites new to the Australian fauna (Acari: Acaridae). Australian Entomologist, 29: 119–122.
Hirschmann, W. 1960. Gangsystematik der Parasitiformes. Teil 3. Die Gattung Dendrolaelaps Halbert 1915. Acarologie. Schriftenreihe für Vergleichende Milbenkunde, 3: 1–27.
Hirschmann, W. 1970. Ruckenhaarbestimmungstabelle der larven von Macrocheles und verwandten gattungen (Geholaspis, Neopodocinum). Teil 77. Acarologie. Schriftenreihe für Vergleichende Milbenkunde, 14: 44–45.
Hirschmann, W. and Wiśniewski, J. 1982. Weltweite Revision der Gattungen Dendrolaelaps Halbert 1915 und Longoseius Chant 1961 (Parasitiformes). Band II. Artenverzeichnisse, Krankheiten, Mißbildungen, Inseminationsapparate, Abbildungen. Acarologie. Schriftenreihe für Vergleichende Milbenkunde, 29: 1–48, pls. I–XIV, 41–94.
Hofstetter, R.W. 2011. Mutualists and phoronts of the southern pine beetle. In: R.N. Coulson and K.D. Klep-
zig (Eds.). Southern Pine Beetle II. Gen. Tech. Rep. SRS-140. U.S. Department of Agriculture Forest Service, Southern Research Station, Asheville, NC, pp. 161–181.
Hofstetter, R.W., Dinkins-Bookwalter, J., Davis, T.S. and Klepzig, K.D. 2015. Chapter 6—Symbiotic Associations of Bark Beetles. In: F.E.V. Vega and R.W. Hofstetter (Eds.). Bark Beetles: Biology and Ecology of Native and Invasive Species. Academic Press, San Diego, pp. 209–245.
Hofstetter, R.W., Moser, J.C. and Blomquist, S.R. 2013. Mites associated with bark beetles and their hyperphoretic ophiostomatoid fungi. Ophiostomatoid Fungi: expanding Frontiers. CBS Biodiversity Series, 12: 165–176.
Hofstetter, R.W., Moser, J.C. and McGuire, R. 2009. Observations on the mite Schizosthetus lyriformis (Acari: Parasitidae) preying on bark beetle eggs and larvae. Entomological News, 120: 397–400.
Houck, M.A. 1994. Adaptation and transition into parasitism from commensalism: a phoretic model. In: M.A. Houck (Ed.). Mites: Ecological and Evolutionary Analyses of Life-history Patterns. Chapman & Hall, New York & London, pp. 252–281.
Houck, M.A. 1999. Phoresy by Hemisarcoptes (Acari: Hemisarcoptidae) on Chilocorus (Coleoptera: Coccinellidae): influence of subelytral ultrastructure. Experimental and Applied Acarology, 23: 97–118.
Houck, M.A. and Cohen, A.C. 1995. The potential role of phoresy in the evolution of parasitism: Radiolabelling (tritium) evidence from an astigmatid mite. Experimental & Applied Acarology, 19: 677–694.
Hughes, A.M. 1976. The Mites of Stored Food and Houses. London, Her Majestyʼs Stationery Office. 1–400 pp.
Hunter, P.E. and Davis, R. 1963. Observations on Histiostoma gordius (Vitz.) (Anoetidae) and other mites associated with Ips beetles. Proceedings of the Entomological Society of Washington, 65: 287–293.
Hurlbutt, H.W. 1967. Digamasellid mites associated with bark beetles and litter in North America. Acarologia, 9: 497–534.
Hurlbutt, H.W. 1971. Gamasellodes bicolor (Berlese, 1918) (Acarina: Ascidae) and its relatives. Acarologia, 12: 476–478.
Hutcheson, H.J., Mertins, J.W., Larson, S.R. and Paskewitz, S. 2015. Acarine hypopi (Sarcoptiformes: Acaridae) on Ixodes scapularis (Ixodida: Ixodidae) in Central North America. Journal of Medical Entomology, 52: 837–841.
Iglesias-Souto, J., Sanchez-Machin, I., Iraola, V., Poza, P., Gonzalez, R. and Matheu, V. 2009. Oral mite anaphylaxis by Thyreophagus entomophagus in a child: a case report. Clinical and Molecular Allergy, 7: 10.
Kadjaia, G.S. 1994. Geographical variability in the Caucasian species of the genus Forcellinia (Acari: Acaridae). Zoosystematica Rossica, 3: 235–236.
Kaliszewski, M., Athias-Binche, F. and Lindquist, E.E. 1995. Parasitism and parasitoidism in Tarsonemina (Acari: Heterostigmata) and evolutionary considerations. In: R.M.J.R. Baker and D. Rollinson (Eds.). Advances in Parasitology. Academic Press, pp. 335–367.
Kalúz, S., Mašán, P. and Moser, J. C. 2003. Morphology and ecology of Schizosthetus simulatrix (Acari, Mesostigmata) associated with galleries of bark beetles (Scolytidae). Biologia (Bratislava), 58: 165–172.
Kamali, K., Ostovan, H. and Atamehr, A. 2001. A Catalog of Mites and Ticks (Acari) of Iran. Islamic Azad University Scientific Publication Center. 1–196 pp.
Karg, W. 1993. Acari (Acarina), Milben. Parasitiformes (Anactinochaeta). Cohors Gamasina Leach: Raubmilben. 2nd edition. Jena, Gustav Fischer Verlag. 1–523 pp.
Kazemi, S. and Rajaei, A. 2013. An annotated checklist of Iranian Mesostigmata (Acari), excluding the family Phytoseiidae. Persian Journal of Acarology, 2: 63–158.
Khaustov, A.A. 1997. [Mites of supercohort Trigynaspida (Parasitiformes. Celaenopsidae, Cercomegistidae) associated with bark beetles (Coleoptera, Scolytidae) of the Crimea]. Biulleten’—Gosudar-
stvennyĭ Nikitskiĭ botanicheskiĭ sad, 78: 63–66. [In Russian]
Khaustov, A.A. 1998. Mites of the genus Pyemotes (Acariformes, Pyemotidae) associated with bark beetles (Coleoptera, Scolytidae) in Crimea. Zoologicheskii Zhurnal, 77: 1419–1424.
Khaustov, A.A. 1999. Redescription of <Pediculoides> ipidarius Redikortsev, 1947, and a description of a new species from the genus Paracarophenax (Acari: Heterostigmata: Acarophenacidae). Acarina, 7: 57–59.
Khaustov, A.A. 2000a. Mites of the family Winterschmidtiidae (Acari, Astigmata), associated with bark beetles (Coleoptera, Scolytidae) in Crimea. Vestnik Zoologii Supplement, 14: 50–59.
Khaustov, A.A. 2000b. Mites of the genus Elattoma (Acariformes, Pygmephoridae) from Crimea and North-West Russia. Vestnik Zoologii, 34: 77–83, 134.
Khaustov, A.A. 2008. Mites of the Family Scutacaridae of Eastern Palaearctic. Akademperiodika, Kiev. 290 pp.
Khaustov, A.A. 2017a. A new species and a new record of mite genus Coleopygmephorus (Acari: Neopygmephoridae) associated with bark beetles (Coleoptera: Curculionidae: Scolytinae) from Russia. International Journal of Acarology, 43: 351–358.
Khaustov, A.A. 2017b. A new species and new records of pygmephoroid mites (Acari: Scutacaridae, Neopygmephoridae, Pygmephoridae) associated with Ips typographus (Coleoptera: Curculionidae: Scolytinae) from Western Siberia, Russia. International Journal of Acarology, 43: 594–602.
Khaustov, A.A. 2017c. A new species of Dolichocybe (Acari: Dolichocybidae) from Western Siberia, Russia. Systematic and Applied Acarology, 22: 1678–1687.
Khaustov, A.A. 2017d. A new species of Tarsonemus (Acari: Tarsonemidae) associated with the bark beetle, Polygraphus proximus (Coleoptera: Curculionidae: Scolytinae) from the Far East of Russia. Acarologia, 57: 673–687.
Khaustov, A.A. 2018. First record of the genus Microdispodides (Acari: Heterostigmata: Pygmephoridae) in Palaearctic, with description of a new species from Western Siberia, Russia. Systematic and Applied Acarology, 23: 441–448.
Khaustov, A.A., Tolstikov, A.V. and Ermilov, S.G. 2017. Parasitoid mites (Acari: Heterostigmata) of bark beetles (Coleoptera: Curculionidae: Scolytinae) of Western Siberia. Abstract book of the third International Persian Congress of Acarology, 23–25 August 2017: 23–24.
Khaustov, A.A., Trach, V.A. and Bobylev, A.N. 2016. Mites (Acari) phoretic on six-toothed spruce bark beetle, Pityogenes chalcographus Linnaeus (Coleoptera: Curculionidae: Scolytinae), in Western Siberia, Russia. Acarina, 24: 137–151.
Kiełczewski, B., Moser, J.C. and Wiśniewski, J. 1983. Surveying the acarofauna associated with Polish Scolytidae. Bulletin de la Société des Amis des Sciences et des Lettres de Poznan, Série D, 22: 151–159.
Kiełczewski, B. and Wiśniewski, J. 1971. Fauna roztoczy na mrówkach w Nadleśnictwie Doświad-
czalnym Zielonka. Prace Komisji Nauk Rolniczych i Komisji Nauk Leśnych, 32: 51–55.
Kiełczewski, B. and Wiśniewski, J. 1980. Bark beetle acarofauna in different types of forest habitat. Part III. Tarsonemini, Prostigmata, Acaridiae. Bulletin de la Société des Amis des Sciences et des Lettres de Poznań, Série D Sciences Biologiques, 20: 161–175.
Kiełczewski, B. and Wiśniewski, J. 1983. Bark beetle acarofauna in different types of forest habitat. I, II. Introduction and Mesostigmata. Folia Forestalia Polonica. Seria A. Leśnictwo, 25: 129–162.
Kinn, D.N. 1967. Notes on the life cycle and habits of Digamasellus quadrisetus (Mesostigmata: Digamasellidae). Annals of the Entomological Society of America, 60: 862–865.
Kinn, D.N. 1968. A new species of Pleuronectocelaeno (Acarina. Celaenopsidae) associated with bark beetles in North and Central America. Acarologia, 10: 191–205.
Kinn, D.N. 1991. A new Pleuronectocelaeno (Acari: Celaenopsidae) from Japan, with remarks on North American and European species. International Journal of Acarology, 17: 103–105.
Kinn, D.N. and Swanston, D.N. 1976. Key to mites commonly associated with the southern pine beetle. New Orleans, LA, Dept. of Agriculture, Forest Service, Southern Forest Experiment Station. 11 pp.
Kirisits, T. 1998. Pathogenicity of three blue-stain fungi associated with the bark beetle Ips typographus to Norway spruce in Austria. Österreichische Zeitschrift für Pilzkunde, 7: 191–201.
Klimov, P.B. 1997. New data on the fauna and biology of acarid mites (Acari, Acaridae) from the Far East. Acarina, 5: 69–72.
Klimov, P.B. 1998a. Review of the genus Boletoglyphus (Acariformes, Acaridae). Zoologicheskii Zhurnal, 77: 1121–1127.
Klimov, P.B. 1998b. To the knowledge of acarid mites of the genus Schwiebea (Acariformes, Acaridae) from the Far East, with notes on systematic of the genus. Vestnik Zoologii, 32: 13–30.
Klimov, P.B. and Khaustov, A.A. 2018. A review of acarid mites (Acariformes: Acaridae) associated with bark beetles (Coleoptera: Curculionidae: Scolytinae), with description of Ipsoglyphus bochkovi gen. and sp. nov. Systematic & Applied Acarology, 23: 1–26.
Klimov, P.B., OConnor, B.M., Ochoa, R., Bauchan, G.R. and Scher, J. 2016. Bee Mite ID: Bee-Associated Mite Genera of the World. USDA APHIS Identification Technology Program (ITP), Fort Collins, CO. Accessed 23 Mar 2018. http://idtools.org/id/mites/beemites/factsheet.php?name=15324
Klimov, P.B. and Tolstikov, A.V. 2011. Acaroid mites of Northern and Eastern Asia (Acari: Acaroidea). Acarina, 19: 252–264.
Knee, W., Forbes, M.R. and Beaulieu, F. 2013. Diversity and host use of mites (Acari: Mesostigmata, Oribatida) phoretic on bark beetles (Coleoptera: Scolytinae): Global generalists, local specialists? Annals of the Entomological Society of America, 106: 339–350.
Kofler, A. and Schmölzer, K. 2000. Zur Kenntnis phoretischer Milben und ihrer Tragwirte in Österreich (Acarina: Gamasina, Uropodina). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 87: 133–157.
Krause, A. 1917 (1919). Eine neue Borkenkäfermilbe, Calvolia Kneissli m. von Orthotomicus laricis Fabr. Archiv für Naturgeschichte. Abteilung A, 83: 123–124.
Kršlak, B., Zach, P. and Kulfan, J. 2010. The role of Hylastes cunicularius Erichson (Coleoptera: Scolytidae) in transferring uropodine mites in a mountain spruce forest. Journal of Forest Science, 56: 258–264.
Kuhlmann, M. 1998. Nachweise mit Bienen und Wespen (Hymenoptera Aculeata) assoziierter Milben (Acari) und Fächerflügler (Strepsiptera). Linzer biologische Beiträge, 30: 69–80.
Kurosa, K. 2000. Mites associated with insects and small mammals in Oda-chô, Ehime Prefecture, southwest Japan. In: Odamiyama-no-Shizen-Henshû-Iinkai and E. Yamamoto (Eds.). Nature of Odamiyama, Oda-chô, Ehime, pp. 897–946.
Laboulbène, A. 1852. M. Al. Laboulbène lit une note conlenant la desription dʼun Acarus qui se développe très souvent sur les insectes, dans nos collections, et auquel il donne le nom dʼA. entomophagus. Annales de la Société entomologique de France, 10: LIV.
Laboulbène, A. and Robin, C. 1862. Description de lʼAcarus (Tyroglyphus) entomophagus Laboulbène et observations anatomiques sur le genre Tyroglyphus. Annales de la Société entomologique de France, 2: 317–338.
Levieux, J., Lieutier, F., Moser, J.C. and Perry, T.J. 1989. Transportation of phytopathogenic fungi by the bark beetle Ips sexdentatus Boerner and associated mites. Journal of Applied Entomology, 108: 1–11.
Li, C., Zhan, X., Sun, E., Zhao, J., Wang, H., He, J. et al. 2014. The density and species of mite breeding in stored products in China. Nutrición Hospitalaria, 31: 798–807.
Li, L.-S., Xuan, J.-Y. and Fan, Q.-H. 1992. Taxonomic investigation of food mites in Sichuan Province. Journal of Southwest Agricultural University, 14: 23–34.
Lieutier, F. 1978. Les acariens associes a Ips typographus et Ips sexdentatus (Coleoptera : Scolytidae) en region parisienne et les variations de leurs populations au cours du cycle annuel. Bulletin dʼécologie, 9: 307–321.
Lignières, J.L.M. 1893. Étude zoologique et anatomique du Tyroglyphus malus et de sa nymphe hypopiale. Mémoires de la Société zoologique de France, 6: 5–15.
Lindquist, E.E. 1969. Review of Holarctic tarsonemid mites (Acarina: Prostigmata) parasitizing eggs of ipine bark beetles. Memoirs of the Entomological Society of Canada, 101: 5–111.
Lindquist, E.E. 1970a. Relationships between mites and insects in forest habitats. Canadian Entomologist, 102: 978–984.
Lindquist, E.E. 1970b. Review of the genus Heterotarsonemus (Acarina: Tarsonemidae). Canadian Entomologist, 102: 812–829.
Lindquist, E.E. 1971. New species of Ascidae (Acarina: Mesostigmata) associated with forest insect pests. Canadian Entomologist, 103: 919–942.
Lindquist, E.E. and Hunter, P.E. 1965. Some mites of the genus Proctolaelaps Berlese (Acarina: Blattisociidae) associated with forest insect pests. Canadian Entomologist, 97: 15–32.
Macfarlane, R.P. 2005. Mites associated with bumble bees (Bombus: Apidae) in New Zealand. Records of the Canterbury Museum, 19: 29–34.
Magowski, W.Ł. 2010. Schaarschmidtia, a new subgenus of the genus Tarsonemus Canestrini and Fanzago, 1876 (Acari: Heterostigmatina: Tarso-
nemidae)—a systematic review of the mite taxon associated with subcortical coleopteran insects, with notes on its host affiliation, ecology and biogeography. Polish Journal of Entomology, 79: i–ii, 1–176.
Magowski, W.Ł. and Khaustov, A.A. 2006. Two new species of Tarsonemus (Acari: Tarsonemidae) associated with bark beetles (Coleoptera: Scolytidae) from Crimea, Ukraine. Zootaxa, 1359: 31–55.
Magowski, W.Ł. and Moser, J.C. 2003. Redescription of Tarsonemus minimax and definition of its species-group in the genus Tarsonemus (Acari: Tarsonemidae) with descriptions of two new species. Annals of the Entomological Society of America, 96: 345–368.
Mahunka, S. 1961. Contributions to the tyroglyphid fauna of Hungary (Acari). Annales Universitatis Scientiarum Budapestinensis de Rolando Eötvös Nominatae. Sectio biologica, 4: 113–117.
Mahunka, S. 1968. Studies on the mite fauna of Hungary I. (Acari). Annales Historico-Naturales Musei Nationalis Hungarici, 60: 249–260.
Mahunka, S. 1970. Atkák V.—Acari V. Magyarország Állatvilága (Fauna Hungariae 101), 18: 1–76.
Mahunka, S. 1976a. New and incompletely known taxa from the families Acaridae and Anoetidae (Acari: Acarida). Acta Zoologica Hungarica, 22: 303–326.
Mahunka, S. 1976b. Über einige an Histeriden (Coleoptera) lebende Anoetiden-Deutonymphen (Acari). Parasitologia Hungarica, 9: 97–104.
Mahunka, S. 1983. Data to the Acarida Fauna of the Hortobágy National Park (Acari). In: S. Mahunka (Ed.), The Fauna of the Hortobágy National Park II. Akadémiai Kiadó, Budapest, pp. 375–376.
Mahunka, S. and Moser, J.C. 1980. Scutacarus scolyti sp.n. a new scutacarid species (Acari: Tarsonemina) from Germany. Parasitologia Hungarica, 13: 99–102.
Mahunka, S. and Moser, J.C. 1982. New data to the knowledge of the tarsonemids (Acari) living on bark beetles. Parasitologia Hungarica, 14: 87–89.
Makarova, O.L. 2012. Gamasid mites (Parasitiformes, Mesostigmata) of the European Arctic and their distribution patterns. Zoologicheskii Zhurnal, 91: 907–927.
Marchenko, I.I. 2002. Faunistic review of free-living Gamasina mites (Acari, Mesostigmata) from Sakhalin and Kuril Islands. Euroasian Entomological Journal: 31–48.
Marchenko, I.I. 2012. Spatial-typological organization of the soil Gamasina mite (Acari, Mesostigmata) community of Northeastern Altai. Communication II. Contemporary Problems of Ecology, 5: 23–33.
Mašán, P. 1999. Mites (Acarina) associated with burying and carrion beetles (Coleoptera, Silphidae) and description of Poecilochirus mrciaki sp. n. (Mesostigmata, Gamasina). Biologia (Bratislava), 54: 515–524.
Mašán, P. 2001. Roztoče kohorty Uropodina (Acarina, Mesostigmata) Slovenska. Bratislava, Slovenské národné múzeum. 320 pp.
Maslov, A.D. 2010. Koroed-Tipograf i Usykhanie Elovykh Lesov. Moscow, Vserosiyskiy nauchno-issledovatelskiy institute lesovodstva I mekhanizatsii lesnogo khozyaystva. 138 pp.
Maslov, A.D. and Matusevich, L.S. 2008. [Mitesʼ (Acari) role in the population dynamics of bark beetles (Coleoptera, Scolytidae)]. Lesnoy Vestnik, 1: 66–72. [In Russian]
Matthewman, W.G. and Pielon, D.P. 1971. Arthropods inhabiting the sporophores of Fomes fomentarius (Polyporaceae) in Gatineau Park, Quebec. Canadian Entomologist, 103: 775–847.
Mazzuco, R.M., Lebarbenchon, M.A., Moecke, E.H.S., Meyer, E., Spíndola-Batti, M.A., Fröde, T. et al. 2000. Anafilaxia por ingestão de ácaros de estocagem. Revista Brasileira de Alergia e Imunopatologia, 23: 189.
Mercado, J.E., Hofstetter, R.W., Reboletti, D.M. and Negron, J.F. 2014. Phoretic symbionts of the Mountain Pine Beetle (Dendroctonus ponderosae Hopkins). Forest Science, 60: 512–526.
Michael, A.D. 1885. Notes on the life-histories of some of the little-known Tyroglyphidae. Journal of the Royal Microscopical Society, 2: 19–32+Pl. III.
Michael, A.D. 1903. British Tyroglyphidae. Vol. 2. London, Ray Society. 1–183+plates i–xxxix. pp.
Michalski, J. and Ratajczak, E. 1989. Korniki (Coleoptera: Scolytidae) wraz z towarzyszącą im fauną w Górach Świętokrzyskich. Fragmenta Faunistica, 32: 279–318.
Moniez, R. 1892. Contribution à lʼhistoire naturelle du Tyroglyphus mycophagus Mégnin. Mémoires de la Société zoologique de France, 5: 584–601.
Moraza, M.L., Fernandez, M. and Jurc, M. 2013. Phoretic mites of the six-spined engraver beetle, Ips sexdentatus (Böerner, 1776) (Coleoptera, Scolytinae), on Pinus halepensis in Slovenia. International Journal of Acarology, 39: 597–599.
Moser, J.C. 1975. Mite predators of southern pine beetle. Annals of the Entomological Society of America, 68: 1113–1116.
Moser, J.C. and Bogenschütz, H. 1984. A key to the mites associated with flying Ips typographus in South Germany. Zeitschrift Fur Angewandte Entomologie-Journal of Applied Entomology, 97: 437–450.
Moser, J.C., Eidmann, H.H. and Regnander, J.R. 1989. The mites associated with Ips typographus in Sweden. Annales Entomologici Fennici, 55: 23–27.
Moser, J.C., Perry, T.J. and Furuta, K. 1997. Phoretic mites and their hyperphoretic fungi associated with flying Ips typographus japonicus Niijima (Col., Scolytidae) in Japan. Journal of Applied Entomology, 121: 425–428.
Moser, J.C., Perry, T.J. and Solheim, H. 1989. Ascospores hyperphoretic on mites associated with Ips typographus. Mycological Research, 93: 513–517.
Moser, J.C. and Roton, L. M. 1971. Mites associated with southern pine bark beetles in Allen Parish, Louisiana. Canadian Entomologist, 103: 1775–1798.
Mullen, G.R., Leprince, D.J. and OConnor, B.M. 1989. Acarine associates of horse flies and deer flies (Diptera: Tabanidae) in North America. In: Progress in Acarology. Volume 2. Oxford & IBH Publishing Co. Pvt. Ltd, New Delhi, pp. 291–296.
Müller, J. 1859 (1860). Insectenepizoen der mährischen Fauna. 157–184.
Musken, H., Franz, J.T., Wahl, R., Paap, A., Cromwell, O., Masuch, G. et al. 2000. Sensitization to different mite species in German farmers: clinical aspects. Journal of Investigational Allergology and Clinical Immunology, 10: 346–351.
Nachev, P.D. and Trenchev, G. 1987. The acarofauna associated with some Coccidae in Bulgaria. Pochvoznanie, Agrokhimiya i Rastitelna Zashchita, 22: 81–86.
Newstead, R. and Morris , H.M. 1920. Bionomic, morphological and economic report on the acarids of stored Grain and Flour—part II. Royal Society Reports on Grain Pests (War) Committee, 8: 4–15+Pl. I–V.
OConnor, B.M. 1982. Evolutionary ecology of astigmatid mites. Annual Review of Entomology: 385–409.
OConnor, B.M. 1989. Systematics, ecology and host associations of Naiadacarus (Acari: Acaridae) in the Great Lakes region. Great Lakes Entomologist, 22: 79–94.
OConnor, B.M. 1991. A preliminary report on the arthropod-associated astigmatid mites (Acari: Acariformes) of the Huron Mountains of Northern Michigan. Michigan Academician, 24: 307–320.
OConnor, B.M. and Houck, M.A. 1989. Two new genera of Hemisarcoptidae (Acari: Astigmata) from the Huron Mountains of northern Michigan. Great Lakes Entomologist, 22: 1–10.
OConnor, B.M. and Pfaffenberger, G.S. 1987. Systematics and evolution of the genus Paraceroglyphus and related taxa (Acari: Acaridae) associated with fleas (Insecta: Siphonaptera). Journal of Parasitology, 73: 1189–1197.
Okabe, K. and OConnor, B.M. 2001. A method for both mass and individual rearing of fungivorous astigmatid mites (Acari). Experimental and Applied Acarology, 25: 493–504.
Olsen, A.R. 1983. Food-contaminating mites from imported foods entering the united states through southern California. International Journal of Acarology, 9: 189–193.
Oshima, S. 1977. Acaroid mites of house dust. In: M. Sa-
sa and J. Aoki (Eds.). Contributions to Acarology in Japan. Zukan-no-Hokuryukan, Tokyo, pp. 525–568.
Ostovan, H., Miri, M. and Beyzavi, G. 2010. The first report of the genus Cercoleipus and family Cercomegistidae (Acari: Mesostigmata) from Iran. Plant Protection Journal, 2: 145–151. [In Persian]
Oudemans, A.C. 1901. Notes on Acari: Third series. Tijdschrift der Nederlandsche Dierkundige Vereeniging. Leiden, 7: 50–87+plates i–iii.
Oudemans, A.C. 1903. Notes on Acari: Sixth Series. Tijdschrift voor Entomologie, 46: 1–24.
Oudemans, A.C. 1906. Acarologische Aanteekeningen XXV. Entomologische Berichten, 2: 121–123.
Oudemans, A.C. 1907. Nachtrag zur Milben-Fauna der Umgegend Bremens. Abhandlungen herausgegeben vom Naturwissenschaftlichen Verein zu Bremen, 19: 48–67+Taf. II (Pope, C.A.: p 47).
Oudemans, A.C. 1919. Acarologische Aanteekeningen XXXVIII. Entomologische Berichten, 3: 183–191.
Oudemans, A.C. 1924. Acarologische Aanteekehingen LXXVII. Entomologische Berichten (Amsterdam), 6: 317–336.
Oudemans, A.C. 1928. Fauna Buruana Acari. Treubia, 7: 37–100.
Oudemans, A.C. 1939. Neue Funde auf dem Gebiete der Systematik und der Nomenclatur der Acari. VII. Zoologischer Anzeiger Leipzig, 127: 184–190.
Penttinen, R., Viiri, H. and Moser, J.C. 2013. The mites (Acari) associated with bark beetles in the Koli national park in Finland. Acarologia, 53: 3–15.
Pernek, M., Hrasovec, B., Matošević, D., Pilaš, I., Kirisits, T. and Moser, J. 2008. Phoretic mites of three bark beetles (Pityokteines spp.) on Silver fir. Journal of Pest Science, 81: 35–42.
Pernek, M., Wirth, S.F., Blomquist, S.R., Avtzis, D.N. and Moser, J.C. 2012. New associations of phoretic mites on Pityokteines curvidens (Coleoptera, Curculionidae, Scolytinae). Central European Journal of Biology, 7: 63–68.
Pfammatter, J.A., Coyle, D.R., Gandhi, K.J.K., Hernandez, N., Hofstetter, R.W., Moser, J.C. et al. 2016. Structure of phoretic mite assemblages across subcortical beetle species at a regional scale. Environmental Entomology, 45: 53–65.
Pfammatter, J.A., Moser, J.C. and Raffa, K.F. 2013. Mites phoretic on Ips pini (Coleoptera: Curculionidae: Scolytinae) in Wisconsin red pine stands. Annals of the Entomological Society of America, 106: 204–213.
Philips, J.R. 2009. The mite (Acarina) fauna of trogid beetles (Coleoptera: Trogidae). International Journal of Acarology, 35: 1–17.
Pielou, D.P. and Verma, A.N. 1968. The arthropod fauna associated with the birch bracket fungus, Polyporus betulinus, in eastern Canada. Canadian Entomologist, 100: 1179–1199.
Plumari, M. 2008. Specie di Celaenopsidae, Parasitidae, Digamasellidae e Ascidae nuove per lʼItalia (Acarina Mesostigmata). Bollettino della Societa Entomologica Italiana, 140: 75–90.
Poliță, D., Manu, M. and Marcu, V.M. 2016. Relationship among phoretic mites and Norway spruce bark beetles—Ips typographus and Pityogenes chalcographus. Revista Pădurilor, 131: 57–65.
Reck, G.F. 1976. Katalog Akarofauny Gruzinskoy SSR [=Catalogue of the acarofauna of the Georgian SSR]. Tbilisi, Metsniereba. 127 pp.
Redikortzev, V.V. 1947. [The mite Pediculoides ipidarius sp. n. (Acari: Pediculoididae), a parasite of bark beetles]. Entomologicheskoe Obozrenie, 29: 247–249. [In Russian]
Ripka, G., Fain, A. and Bolland, H.R. 1999. New data to the knowledge on the corticolous mite fauna in Hungary (Acari: Prostigmata, Astigmata, Oribatida). Acta Phytopathologica et Entomologica Hungarica, 34: 363–371.
Ripka, G. and Szabo, A. 2010. Additional data to the knowledge of the mite fauna of Hungary (Acari: Mesostigmata, Prostigmata and Astigmata). Acta Phytopathologica et Entomologica Hungarica, 45: 373–381.
Salmane, I. 2001. A check-list of Latvian Gamasina mites (Acari, Mesostigmata) with short notes to their ecology. Latvijas Entomologs, 38: 27–38.
Salmane, I. 2005. List of Mesostigmata mites (Acari, Parasitiformes) associated with Aphyllophorales fungi (Basidiomycetes) in Latvia. Latvijas Entomologs, 42: 45–57.
Salmane, I. 2006. New Mesostigmata (Acari, Parasitiformes) species in the fauna of Latvia. Latvijas Entomologs, 43: 52–56.
Salmane, I. 2007. Mesostigmata mite (Acari, Parasitiformes) fauna of wood-related microhabitats in Latvia. Latvijas entomologs, 44: 69–86.
Salmane, I. and Telnov, D. 2009. Mesostigmata mites (Acari: Parasitiformes) associated with beetles (Insecta: Coleoptera) in Latvia. Latvijas Entomologs, 47: 58–70.
Samšiňák, K. 1956. Roztoci (Acari) na mouse Laphria flava L. Věstník Československé společnosti zoolo-
gické, 20: 353–357.
Samšiňák, K. 1957. Einige Bemerkungen zur Faunistik der in Gesellschaft von Insekten lebenden Acari. Acta faunistica entomologica Musei Nationalis Pragae, 2: 109–114.
Samšiňák, K. 1958. Einige neue Acariden-Deutonymphen als Commensalen der Insekten (Acari). Časopis Československé společnosti entomologické, 55: 289–295.
Samšiňák, K. 1960. Über einige forstwirtschaftlich wichtige Milben der Gattung Proctolaelaps Berlese 1923. Československá Parasilogie, 7: 297–307.
Samšiňák, K. 1962. Beiträge zur Kenntnis der Familie Anoetidae (Acari). Časopis Československé společnosti entomologické, 59: 87–97.
Samšiňák, K. and Vobrázková, E. 1983. Mites from the city pavement. Věstnik Československé Zoologické Společnosti, 47: 118–121.
Schaarschmidt, L. 1959. Systematik und Ökologie der Tarsonemiden. In: H.-J. Stammer (Ed.). Beiträge zur Systematik und Ökologie mitteleuropäischer Acarina. Vol. 1, part 2. Akademische verlagsgesellschaft Geest, Portig K.G., Leipzig, pp. 713–823.
Scheucher, R. 1957. Systematik und Ökologie der deutschen Anoetinen. In: H.-J. Stammer (Ed.). Beiträge zur Systematik und Ökologie mitteleuropäischer Acarina. Vol. 1, pp. 233–384.
Schweizer, J. 1949. Die Landmilben des schweizerischen Nationalparkes. 1. Teil. Parasitiformes Reuter 1909. Ergebnisse der wissenschaftlichen Untersuchungen im Schweizerischen Nationalpark. Heft 21. Band || (Neue Folge). Lüdin, Liestal. 1–99 pp.
Sellnick, M. 1951. Pleuronectocelaeno austriaca (Vitzthum) und P. africana sp. nov. (Acari). Entomologisk Tidskrift, 72: 184–194.
Sevastianov, V.D. 1969. A new myrmecophilous mite Divilia oculata gen. et sp.n. (Sarcoptiformes, Saproglyphidae). Zoologicheskii Zhurnal, 48: 447–449.
Sevastianov, V.D. 1970. Acarofauna of the forest ant Formica rufa L. In: I.S. Eytminavichyute (Ed.). Oribatidy i ikh rol’ v pochvoobrazovatelnykh protsessakh. Institute of Zoology and Parasitology, Lithuanian SSR Academy of Sciences, Vilnius pp. 143–147.
Sevastianov, V.D. 1983. New species of the genus Scutacarus (Trombidiformes, Scutaridae). Zoologicheskii Zhurnal, 62: 1487–1496.
Shcherbak, G.I. 1980. Kleshchi semeystva Rhodacaridae Palearktiki [Mites of the family Rhodacaridae of the Palaearctic]. Naukova Dumka, Kiev. 215 pp.
Shcherbak, G.I. and Balan, P.G. 1982. Gamasid mites from the sand martin nests of the Central Forest Steppe of the Ukrainian SSR. Vestnik Zoologii, 1982: 12–19. [In Russian]
Simoni, S., Nannelli, R., Roversi, P.F., Turchetti, T. and Bouneb, M. 2014. Thyreophagus corticalis as a vector of hypovirulence in Cryphonectria parasitica in chestnut stands. Experimental and Applied Acarology, 62: 363–375.
Sinha, R.N. 1964. Mites of stored grain in Western Canada—ecology and methods of survey. Proceedings of the Entomological Society of Manitoba, 20: 19–33.
Skorupski, M. and Gwiazdowicz, D.J. 1996. Roztocze (Acari, Mesostigmata) Pienin. Fragmenta Faunistica (Warsaw), 39: 223–243.
Smiley, R.L. 1969. Further studies on the Tarsonemidae 2. (Acarina). Proceedings of the Entomological Society of Washington, 71: 218–229.
Smiley, R.L. and Moser, J.C. 1970. Three cheyletids found with pine bark beetles (Acarina: Cheyletidae). Proceedings of the Entomological Society of Washington, 72: 229–235.
Smiley, R.L. and Moser, J.C. 1985. A new species, key to females, and distribution records for Heterotarsonemus (Acari: Tarsonemidae). International Journal of Acarology, 11: 247–253.
Stephen, F.M. and Kinn, D.N. 1980. Spatial distribution of mite associates of within-tree populations of Dendroctonus frontalis Zimm. Environmental Entomology, 9: 713–715.
Summers, F.M. and Price, D.W. 1970. Review of the mite family Cheyletidae. University of California Publications in Entomology, 61: 1–153.
Takov, D., Pilarska, D. and Moser, J. 2009. Phoretic mites associated with spruce bark beetle Ips typographus L. (Curculionidae: Scolytinae) from Bulgaria. Acta Zoologica Bulgarica, 61: 293–296.
Tichomirov, S.I. 1977. Family Parasitidae. In: M.S. Gilarov and N.G. Bregetova (Eds.). Opredelitel’ obytayshchikh v pochve kleshchey Mesostigmata [Identification key to soil-inhabiting mites Mesostigmata]. Nauka, Leningrad, pp. 55–107. [In Russian]
Trach, V.A. and Khaustov, A.A. 2017a. Redescription of adults and description of juvenile stages of bark beetle-associated mite Cercoleipus kuznetsovi Khaustov, 1997 (Acari: Mesostigmata: Cercomegistidae). Systematic and Applied Acarology, 22: 1733–1747.
Trach, V.A. and Khaustov, A.A. 2017b. Mites of the genus Proctolaelaps Berlese, 1923 (Acari: Mesostigmata: Melicharidae) associated with bark beetles in Asian Russia. Acarina, 25: 151–163.
Tseng, Y.H. 1989. Redescription of Tyrophagus kentinus Tseng, Tyrophagus bambusae Tseng and Aleuroglyphagus formosanus Tseng and describes a new record species Rhizoglyphus canadii Manson. Chinese Journal of Entomology, Special Publication, 3: 37–50.
Türk, E. and Türk, F. 1957. Systematik und Ökologie der Tyroglyphiden Mitteleuropas. In: H.-J. Stammer (Ed.). Beiträge zur Systematik und Ökologie mitteleuropäischer Acarina. Band 1. Tyroglyphidae und Tarsonemini. Akademische Verlagsgesellschaft Geest & Portig K.-G., Leipzig, pp. 4–384.
Turk, F.A. and Turk, S.M. 1952. LV.—Studies of Acari.—7th series: “Records and descriptions of mites new to the British fauna, together with short notes on the biology of sundry species”. Annals and Magazine of Natural History, 5: 475–506.
Vissa, S. and Hofstetter, R.W. 2017. The role of mites in bark and ambrosia beetle-fungal interactions. In: V. Shields (Ed.). Insect Physiology and Ecology. InTech, Rijeka, pp. 135–156.
Vitzthum, H. 1926. Acari als Commensalen von Ipiden (Der Acarologischen Beobachtungen 11. Reihe). Zoologische Jahrbücher. Abteilung für Systematik, Ökologie und Geographie der Tiere, 52: 407–503.
Volgin, V.I. 1953. [Two new species of acaroid mites (Acarina, Acaroidea) of the fauna of the USSR]. Entomologicheskoe Obozrenie, 33: 262–265.
Volgin, V.I. 1965. [Two new species of acaroid mites (Acarina, Acaroidea) of the fauna of the USSR]. Entomologicheskoe Obozrenie, 44: 203–211.
Vrabec, M., Kalúz, S. and Ferenčík, J. 2012. Foretické roztoče na lykožrútovi smrekovom (Ips typographus) na vybraných lokalitách vo Vysokých Tatrách. Entomofauna Carpathica, 24: 1–14.
Walter, D.E. 1987. Life history, trophic behavior, and description of Gamasellodes vermivorax n.sp. (Mesostigmata: Ascidae), a predator of nematodes and arthropods in semiarid grassland soils. Canadian Journal of Zoology, 65: 1689–1695.
Walter, D.E. and Krantz, G.W. 2009. Collection, rearing, and preparing specimens. In: G.W. Krantz and D.E. Walter (Eds.). A Manual of Acarology. Third Edition. Lubbock (Texas), Texas Tech University Press, pp. 83–96.
Walter, D.E. and Latonas, S. 2012. Almanac of Alberta Acari Part II. Ver. 2.1. Edmonton, AB, The Royal Alberta Museum. 159 pp. http://www.royalalbertamuseum.ca/natural/insects/research/research.htm
Wasmann, E. 1897. Über einige myrmecophile Acarinen. I. Zoologischer Anzeiger, 20: 170–173.
Wasylik, A. 1959. Mite fauna (Tyroglyphoidea) in the nests of the common sparrow (Passer domesticus L.). Ekologia Polska. Seria B, 5: 187–190.
Westerboer, I. 1963. Die Familie Podocinidae Berlese 1916. In: H.-J. Stammer (Ed.). Beiträge zur Systematik und Ökologie Mitteleuropäischer Acarina. Band II. Mesostigmata 1. Akademische Verlagsgesellschaft Geest & Portig K.-G., Leipzig, pp. 179–450.
Wirth, S.F. 2016. Description of developmental instars of Bonomoia sibirica sp.n. (Astigmata: Histiostomatidae) with ecological observations and phylogenetic conclusions. Acarina, 24: 97–111.
Wirth, S.F. and Garonna, A.P. 2015. Histiostoma ovalis (Histiostomatidae, Acari) associated with Ips sexdentatus (Scolytinae, Curculionidae, Coleoptera): ecology and mite redescription on the basis of formerly unknown adults and nymphs. International Journal of Acarology, 41: 415–428.
Wirth, S.F. and Pernek, M. 2012. First record of the mite Histiostoma ulmi in silver fir and indication of a possible phoretic dispersal by the longhorn beetle Acanthocinus reticulatus. Šumarski List, 136: 597–603.
Wirth, S.F., Weis, O. and Pernek, M. 2016. Comparison of phoretic mites associated with bark beetles Ips typographus and Ips cembrae from Central Croatia. Šumarski List, 140: 549–560.
Womersley, H. 1941. Studies in Australian Acarina. (2) Tyroglyphidae (s. l.). Records of the South Australian Museum, 6: 451–488.
Woodring, J.P. and Moser, J.C. 1970. Six new species of anoetid mites associated with North American Scolytidae. The Canadian Entomologist, 102: 1237–1257.
Woodroffe, G.E. 1953. An ecological study of the insects and mites in the nests of certain birds in Britain. Bulletin of Entomological Research London, 44: 739–772.
Wurst, E. 2002. Beitrag zur Revision der mitteleuropäischen Arten der Gattung Schwiebea Oudemans, 1916 (Acari: Acaridae). Verlag Grauer, Beuren and Stuttgart. 128 pp. + plates.
Wurst, E. 2006. Limnic Acaridida (Astigmata). In: R. Ge-
recke (Ed.). Süßwasserfauna von Mitteleuropa, Vol. 7/2-1 Chelicerata: Araneae/Acari I. Süßwasserfauna von Mitteleuropa. Springer Spektrum, Berlin and Heidelberg, pp. 38–88.
Wurst, E. and Frank, B. 1998. Contributions to a revision of the genus Schwiebea (Acari: Acaridae). 1. Redescription of Schwiebea talpa and Schwiebea nesbitti. Stuttgarter Beitraege zur Naturkunde Serie A (Biologie), 579: 1–52.
Zach, P., Kršiak, B., Kulfan, J., Parák, M. and Kontschán, J. 2016. Mites Trichouropoda and Uroobovella spp. (Uropodoidea) phoretic on bark beetles (Scolytinae): a comparison from a declining mountain spruce forest in Central Europe. International Journal of Acarology, 42: 212–217.
Zamec, R. 2013. Soil astigmatine (Acari: Astigmatina) mites from birdsʼ nests in Slovakia. Acta Societatis Zoologicae Bohemicae, 77: 165–172
Zachvatkin, A.A. 1941. Tiroglifoidnye kleshchi Tyroglyphoidea [=Tyroglyphoid mites Tyroglyphoidea]. In: S.A. Zernov (Ed). Fauna SSSR: Paukoobraznuye. Akademiya Nauk SSSR, Moscow–Leningrad, p. 475.
Ždárková, E. 1967. Stored food mites in Czechoslovakia. Journal of Stored Products Research, 3: 155–175.